A new species of killifish of the family Profundulidae from the highlands of the Mixteca region, Mexico

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"autoresLista" => "Claudia Patricia Ornelas-García, Emilio Martínez-Ramírez, Ignacio Doadrio" "autores" => array:3 [ 0 => array:2 [ "nombre" => "Claudia Patricia" "apellidos" => "Ornelas-García" ] 1 => array:2 [ "nombre" => "Emilio" "apellidos" => "Martínez-Ramírez" ] 2 => array:2 [ "nombre" => "Ignacio" "apellidos" => "Doadrio" ] ] ] ] ] "idiomaDefecto" => "en" "EPUB" => "https://multimedia.elsevier.es/PublicationsMultimediaV1/item/epub/S1870345316301439?idApp=UINPBA00004N" "url" => "/18703453/0000008700000004/v3_201612090152/S1870345316301439/v3_201612090152/en/main.assets" ] ] "en" => array:21 [ "idiomaDefecto" => true "cabecera" => "Taxonomy and systematics" "titulo" => "A new species of killifish of the family Profundulidae from the highlands of the Mixteca region, Mexico" "tieneTextoCompleto" => true "paginas" => array:1 [ 0 => array:2 [ "paginaInicial" => "926" "paginaFinal" => "933" ] ] "autores" => array:1 [ 0 => array:4 [ "autoresLista" => "Claudia Patricia Ornelas-García, Emilio Martínez-Ramírez, Ignacio Doadrio" "autores" => array:3 [ 0 => array:4 [ "nombre" => "Claudia Patricia" "apellidos" => "Ornelas-García" "email" => array:1 [ 0 => "patriciaornelasg@gmail.com" ] "referencia" => array:3 [ 0 => array:2 [ "etiqueta" => "a" "identificador" => "aff0005" ] 1 => array:2 [ "etiqueta" => "b" "identificador" => "aff0010" ] 2 => array:2 [ "etiqueta" => "*" "identificador" => "cor0005" ] ] ] 1 => array:3 [ "nombre" => "Emilio" "apellidos" => "Martínez-Ramírez" "referencia" => array:1 [ 0 => array:2 [ "etiqueta" => "c" "identificador" => "aff0015" ] ] ] 2 => array:3 [ "nombre" => "Ignacio" "apellidos" => "Doadrio" "referencia" => array:1 [ 0 => array:2 [ "etiqueta" => "d" "identificador" => "aff0020" ] ] ] ] "afiliaciones" => array:4 [ 0 => array:3 [ "entidad" => "Departamento de Zoología, Facultad de Ciencias Naturales, Universidad Autónoma de Querétaro, Av. de las Ciencias s/n, 76230 Juriquilla, Querétaro, Mexico" "etiqueta" => "a" "identificador" => "aff0005" ] 1 => array:3 [ "entidad" => "Departamento de Zoología, Universidad Nacional Autónoma de México, Trecer Circuito s/n, Ciudad Universitaria, Coyoacán, 04510 Mexico City, Mexico" "etiqueta" => "b" "identificador" => "aff0010" ] 2 => array:3 [ "entidad" => "Departamento de Investigación, Centro Interdisciplinario de Investigación para el Desarrollo Integral Regional Unidad Oaxaca, Instituto Politécnico Nacional, Calle Hornos N° 1003, Municipio Santa Cruz Xoxocotlán, 71230 Oaxaca, Mexico" "etiqueta" => "c" "identificador" => "aff0015" ] 3 => array:3 [ "entidad" => "Departamento de Biodiversidad y Biología Evolutiva, Museo Nacional de Ciencias Naturales, CSIC, José Gutiérrez Abascal 2, 28006 Madrid, Spain" "etiqueta" => "d" "identificador" => "aff0020" ] ] "correspondencia" => array:1 [ 0 => array:3 [ "identificador" => "cor0005" "etiqueta" => "⁎" "correspondencia" => "Corresponding author." ] ] ] ] "titulosAlternativos" => array:1 [ "es" => array:1 [ "titulo" => "Una especie nueva de ciprinodontiforme de la familia Profundulidae de las tierras altas de la región Mixteca, México" ] ] "resumenGrafico" => array:2 [ "original" => 0 "multimedia" => array:7 [ "identificador" => "fig0020" "etiqueta" => "Figure 4" "tipo" => "MULTIMEDIAFIGURA" "mostrarFloat" => true "mostrarDisplay" => false "figura" => array:1 [ 0 => array:4 [ "imagen" => "gr4.jpeg" "Alto" => 2486 "Ancho" => 2988 "Tamanyo" => 248813 ] ] "descripcion" => array:1 [ "en" => "Results from the discriminant function analysis (DFA): (a) proportions of correctly assigned individuals: 97% for P. mixtlanensis sp. nov. (individuals 1–35 in the graph) and 100% for P. oaxacae (individuals 36–50); (b) distribution frequencies of root factor from the DFA between species." ] ] ] "textoCompleto" => "IntroductionThe family Profundulidae is one of few families of freshwater fishes endemic to the Mesoamerican region. It has been suggested that its history goes back to the Pliocene, or even the Miocene (<a class="elsevierStyleCrossRef" href="#bib0030">Miller, 1955</a>). Currently, most species in the family exhibit low abundances, and their distribution is restricted to the headwaters and spring-fed-tributaries of rivers in southern Mexico, El Salvador, Honduras and Guatemala, from both Pacific and Atlantic versants. Species diversity in the family is fairly low, consisting of 8 species in 2 genera: Profundulus and Tlaloc (<a class="elsevierStyleCrossRef" href="#bib0045">Morcillo, Ornelas-García, Alcaraz, Matamoros, & Doadrio, 2015</a>). In the genus Tlaloc, 4 species are currently recognized: T. candalarius, T. hildebrandi, T. labialis and T. portillorum, all of which exhibit limited distributions. The genus Profundulus also has 4 species, namely P. punctatus, P. oaxacae, P. guatemalensis and P. kreiseri, but has a decidedly wider distribution range.In an early study dealing with systematics of Profundulidae, <a class="elsevierStyleCrossRef" href="#bib0030">Miller (1955)</a> pointed out a wide degree of morphological variation, being more evident within the P. punctatus “group”, in which more than one species might be recognized. Years later, within the P. punctatus “group” <a class="elsevierStyleCrossRef" href="#bib0035">Miller (2005)</a> corroborated the validity of P. oaxacae (<a class="elsevierStyleCrossRef" href="#bib0025">Meek, 1902</a>), whose distribution was restricted to altitudes above 1,550m in the Río Atoyac – Verde Basin, Oaxaca, in southern Mexico (<a class="elsevierStyleCrossRef" href="#bib0020">Martínez-Ramírez, Doadrio-Villarejo, & de Sostoa-Fernández, 2004</a>). Furthermore, in a genetic population analysis of the subgenus Profundulus based on allozymes (<a class="elsevierStyleCrossRef" href="#bib0010">Doadrio, Carmona, Martínez, & De Sostoa, 1999</a>), at least 5 different lineages were recovered. Of these, 2 distinct lineages were found in the Mixteca region and were well differentiated from P. punctatus. One of them corresponded to P. oaxacae and the other one included a clade named P. sp. nov. 1, which corresponded to populations from Mixteca headwaters (with an elevation over 700m) and showed a high degree of molecular divergence from the rest of species of the subgenus Profundulus (Nei's distances D=0.165−0.845). In fact, based on the aforementioned analysis, P. sp. nov. 1 was recovered as a sister clade to the rest of species in the P. punctatus “group”, where P. oaxacae is included.As a consequence of the striking morphological differences between P. oaxacae and P. punctatus, Miller recognized the former as a valid species (<a class="elsevierStyleCrossRef" href="#bib0035">Miller, 2005</a>). These differences include the number of scales around the body (25–28 in P. punctatus vs. 30 in P. oaxacae), and the lateral color pattern (prominent round spots in P. punctatus vs. narrow weak and vertical marks in P. oaxacae).Another aspect clearly separating this pair of species is their distribution, in which P. oaxacae is restricted to the upper part of the río Verde basin, Oaxaca while P. punctatus inhabits the lower part of the same basin. P. sp. nov. 1 also inhabits the upper part of the río Verde basin, but shows a high genetic distinctiveness regarding both, P. oaxacae and P. punctatus (<a class="elsevierStyleCrossRef" href="#bib0010">Doadrio et al., 1999</a>). Consequently, in this study we evaluated the morphological distinctiveness between P. oaxacae and P. sp. nov. 1, and determined whether morphological variation corresponds to the genetic differentiation previously documented by <a class="elsevierStyleCrossRef" href="#bib0010">Doadrio et al. (1999)</a>.Materials and methodsSample collection and taxonomic determinationA total of 5 localities within the Mixteca region were surveyed using cast nets and electrofishing (<a class="elsevierStyleCrossRef" href="#fig0005">Fig. 1</a>). A total of 50 specimens were collected, with a range of individuals per locality from 5 to 15 (<a class="elsevierStyleCrossRef" href="#tbl0005">Table 1</a>). Fin clips were obtained and individuals tagged. Fin clips and voucher specimens were preserved in 95% ethanol and deposited in Colección Nacional de Peces, Instituto de Biología, Universidad Nacional Autónoma de México (CNPE-IBUNAM), Colección de Peces, El Colegio de la Frontera Sur (ECOSC), Museo Nacional de Ciencias Naturales de Madrid, España (MNCN) and Colección de Peces del Centro de México Dr. Edmundo Díaz Pardo, Universidad Autónoma de Querétaro, Mexico (CPCMEDP-UAQ). Localities for P. sp. nov. 1 were determined based on a previous molecular study (<a class="elsevierStyleCrossRef" href="#tbl0005">Table 1</a>; <a class="elsevierStyleCrossRef" href="#bib0010">Doadrio et al., 1999</a>). For P. oaxacae, we measured 15 specimens from Rio Chiquito, Tlaxiaco, in the Mixteco-Balsas basin. P. oaxacae were identified using Miller's keys (<a class="elsevierStyleCrossRef" href="#bib0035">Miller, 2005</a>).<elsevierMultimedia ident="fig0005"></elsevierMultimedia><elsevierMultimedia ident="tbl0005"></elsevierMultimedia>Morphometric analysesFor comparison between species groups, we performed 17 morphometric measurements and 5 meristic counts in 50 individuals from 5 localities (<a class="elsevierStyleCrossRef" href="#fig0005">Fig. 1</a>, <a class="elsevierStyleCrossRefs" href="#tbl0005">Tables 1 and 2</a>). In order to identify the variables that contributed the most to the variation among populations, we performed principal component analysis (PCA) on the morphometric dataset, based on a covariance matrix. Finally, we performed discriminant function analysis (DFA) and linear discriminant analysis (LDA) in order to evaluate each specimen, assign it to a species and assess the degree of separation between the groups.<elsevierMultimedia ident="tbl0010"></elsevierMultimedia>For both analyses, measurements were standardized using the general allometric equation: Ms=Mo (Ls/Lo)b, where Ms=standardized measurement, Mo=measured character length (mm), Ls=overall (arithmetic) mean of the standard component (standard length – SL or head length – HL) for all individuals of each taxon, Lo=standard component of the specimen. For each character “b” was estimated using the non-linear equation, M=aLb as the slope of the regression of logMo on logLo (<a class="elsevierStyleCrossRefs" href="#bib0015">Elliot, Haskard, & Koslow, 1995; Ruiz-Campos, Camarena-Rosales, Varela-Romero, Sánchez-Gonzáles, & De La Rosa-Vélezv, 2003</a>) (logistic regression in Supplementary material 1). Abbreviations for morphological measurements are given in <a class="elsevierStyleCrossRef" href="#tbl0010">Table 2</a>.Two-way analysis of variance (ANOVA) was carried out considering sex and species as factors to evaluate the influence of sexual dimorphism, and the sex×species interaction was included to test if dimorphism varied significantly between species groups for the 17 morphometric measurements. Proportions between measurements were calculated in relation to SL and HL (<a class="elsevierStyleCrossRef" href="#tbl0015">Table 3</a>). Frequencies of the 5 meristic variables were compared between species groups. Mann–Whitney U tests were carried out on the 5 meristic variables as well as on the morphometric proportion measurements. Statistica v.6.0 (<a class="elsevierStyleCrossRef" href="#bib0055">Statistica, 2001</a>) was used for all statistical analyses.<elsevierMultimedia ident="tbl0015"></elsevierMultimedia>Diagnostic measurements were expressed as percent of the standard length (SL) (<a class="elsevierStyleCrossRef" href="#bib0005">Álvarez del Villar, 1970</a>). Additionally, the diagnosis was completed with other relevant features regarding morphology (i.e., body profile, fin shapes, among others), pigmentation and ecology.DescriptionProfundulus mixtlanesis Ornelas-García, Martínez & Doadrio, new species (<a class="elsevierStyleCrossRef" href="#fig0010">Fig. 2</a>).<elsevierMultimedia ident="fig0010"></elsevierMultimedia>Type locality: Spring in Santiago Yosondúa, Oaxaca, MexicoAs a member of the Profundulus subgenus this species shows a preorbital region nearly covered with well-developed scales, a humeral spot, and a densely scaled basal half of the caudal fin. P. mixtlanensis sp. nov. differs from other species in the genus by having a pronounced lower jaw, which markedly differentiates it from P. oaxacae and the remaining species within the P. punctatus “group” sensu<a class="elsevierStyleCrossRef" href="#bib0030">Miller (1955)</a>. Another striking difference between the new species and other species in the subgenus, especially P. oaxacae, is the absence of dark body spotting on the posterior half of the body caused by melanization of the base of the scales. In lateral view, the dorsal profile of the head shows a concavity just above the supra-occipital crest, which quickly reaches a flat profile posteriorly, an attribute that becomes more pronounced with age. In general terms, P. mixtlanesis depicts a longer body and has a lower height than the other species in the subgenus (see <a class="elsevierStyleCrossRef" href="#tbl0015">Table 3</a>). Body height is <2/3 of SL, and pre-pectoral distance <1/3 of SL, in contrast the P. oaxacae proportions are significantly higher reflecting larger body height and pre-pectoral distance for the P. oaxacae. The distance between ventral and anal fins is 15% of SL, lower than that of P. oaxacae. Caudal peduncle depth is less than in P. oaxacae, being 13% of SL (15% of SL in P. oaxacae). The postanal length is 15% of SL, larger than the rest of species (data not shown). The head is shorter than in P. oaxacae, 26–28% of SL in contrast to 28–30% of SL in P. mixtlanensis. Preorbital length, 26–30% of HL, is longer than in P. oaxacae. Eye diameter is smaller than in P. oaxacae, 26–28% of SL. The anal fin is longer than in P. oaxacae, 16–17% of SL. Among meristic characteristics, the following are diagnostic: 30–35 lateral scales, 12–21 rays in the anal fin, and 14–17 rays in the pectoral fin.Taxonomic summaryDisposition of types. Holotype CNPE-IBUNAM 19112 (Female, T1339) and allotype CNPE-IBUNAM 19113 (male, T1340), Spring system in Santiago Yosondúa, Oaxaca, México. The system is composed of five small springs, ∼1m in diameter and less than 1m deep each. Deposited at the CNPE-IBUNAM.Paratypes. (45), Spring, Aldama, Oaxaca (15, CPCMEDP-UAQ:001/T1317-31); río Cucharas, El Carmen Tuxtitlán, Putla de Guerrero, Oaxaca (3, CPCMEDP-UAQ:002/1332-1335); Spring, Santiago Yosondúa, Oaxaca (6, 3 CNPE-IBUNAM 19114, 3; CPCMEDP-UAQ:003/T1340-1346-); río Grande, Santiago Juxtlahuaca, Oaxaca (3, ECOSC 7562-1/ECOSC 7562-3 and 2, CPCMEDP-UAQ:004/T1350-51, CPCMEDP-UAQ). Río Cucharas (10; MNCN:131746-53).Distribution. Springs of the Pacific slope of the Mixteca region of Oaxaca, Mexico in the following river systems: Colorado River, Atoyaquillo River, the Atoyac river basin, and the Mixteco river basin (<a class="elsevierStyleCrossRef" href="#fig0005">Fig. 1</a>). Most of the populations for the species were found above 2,000m, however, they occur at altitudes between 710 and 2,300m.Etymology. The epithet “mixtlanensis” derives from the word “Mixtlan” in the Nahuatl language. The Tenoch culture used to call Mixtlan the region where the new species is found. Mixtlan is a compound word, which is made up of the words Mixtli (cloud) and -Tlan (place). Hence it means “the place of the clouds” (<a class="elsevierStyleCrossRef" href="#bib0040">Montemayor, 2008</a>).ResultsMorphometric analyses showed a separation between the two species groups tested. The two-way ANOVA using 17 morphometric measurements showed significant differences between species for most of the comparisons (p≤0.001, <a class="elsevierStyleCrossRef" href="#tbl0010">Table 2</a>). Analyses however, did not disclose any significant differences in sex (p>0.05), or sex×species interaction (p>0.05, data not shown). Because results did not support the existence of sexual dimorphism within or between species, all posterior analyses were performed without sex separations. Ordination using PCA allowed for identification of two separate clusters of points. Each cluster corresponded to P. oaxacae and to P. sp. nov. 1 (<a class="elsevierStyleCrossRef" href="#fig0015">Fig. 3</a>). PCI explained 91.73% of the variance, where the most important variables in the formation of the groups were SL, PrOL and ED in the positive side of PCI, and PrPD, PrPA and PrPV in the negative side. PCII explained 2.79% of the variance, in which the most important variables were PrPD, PrPA, PrPP and BD, whereas PCIII explained 1.70% of the variance, with the most important variables PrPP, HL, PVL and VAL (Eigenvalues in Supplementary material 4).<elsevierMultimedia ident="fig0015"></elsevierMultimedia>DFA also showed significant discrimination between species. All individuals assigned to P. oaxacae were correctly classified whereas assignation to P. sp. nov. 1 was 97.14% correct (<a class="elsevierStyleCrossRef" href="#fig0020">Fig. 4</a>a, F(16,33)=7.0769 p<0.0001; posterior probabilities are shown in Supplementary materials 2 and 3). Linear discriminant factor from the DFA showed a contrasting distribution between the two groups of cases (<a class="elsevierStyleCrossRef" href="#fig0020">Fig. 4</a>). The root factor was ordered mostly according to the following variables: PrPD and AFL on the positive side and PrPA, DFL and BLD on the negative side.<elsevierMultimedia ident="fig0020"></elsevierMultimedia>Results of the Mann–Whitney U test applied to meristic variables were also significant for three of five variables tested between species (<a class="elsevierStyleCrossRef" href="#tbl0010">Table 2</a>), and when applied to proportions, comparisons were significant for 10 of 16 tested (<a class="elsevierStyleCrossRef" href="#tbl0015">Table 3</a>). Highly significant differences were found in the following proportions: PrPA/SL, PrPP/SL, BD/SL, ADL/SL, BLD/SL, APL/SL, HL/SL, PrOL/HL and ED/SL.Even when there is slight overlap, significant differences were found between species in the following meristic variables (<a class="elsevierStyleCrossRef" href="#tbl0015">Table 3</a>; p<0.001): number of lateral scales, with a mean of 33 and range of 30–35 in P. mixtlanensis, and 31 mean and range of 29–32 in P. oaxacae; and with a lower number of anal-fin rays for P. oaxacae (12–14), while for P. mixtlanensis the anal-fin rays range from 12 to 21 (<a class="elsevierStyleCrossRef" href="#fig0025">Fig. 5</a>). Given the previous results, we conclude that P. mixtlanensis fulfills the criteria to be considered a different taxonomic unit, using both morphological and molecular (<a class="elsevierStyleCrossRef" href="#bib0010">Doadrio et al., 1999</a>) criterion.<elsevierMultimedia ident="fig0025"></elsevierMultimedia>DiscussionP. mixtlanensis and P. oaxacae occur at geographically distant locations in the Mixteco basin, and as we observed in the present description, the two species are easily distinguishable by their body shapes and coloration patterns. P. oaxacae shows a characteristic spotty pattern along the lateral part of the body while in P. mixtlanensis only shows a humeral spot. The molecular distinctiveness of P. mixtlanensis with respect to the other species within the Profundulus genus has been demonstrated by an allozyme study (<a class="elsevierStyleCrossRef" href="#bib0010">Doadrio et al., 1999</a>). In addition, phylogenetic analysis carried out by the authors showed that both, P. oaxacae and P. mixtlanensis depict a high degree of divergence from the rest of the species within the subgenus Profundulus (Dp=10.5±5.9 mtDNA and Dp=1.7±0.2 nDNA, respectively, data not shown)." "textoCompletoSecciones" => array:1 [ "secciones" => array:11 [ 0 => array:3 [ "identificador" => "xres582961" "titulo" => "Abstract" "secciones" => array:1 [ 0 => array:1 [ "identificador" => "abst0005" ] ] ] 1 => array:2 [ "identificador" => "xpalclavsec599352" "titulo" => "Keywords" ] 2 => array:3 [ "identificador" => "xres582960" "titulo" => "Resumen" "secciones" => array:1 [ 0 => array:1 [ "identificador" => "abst0010" ] ] ] 3 => array:2 [ "identificador" => "xpalclavsec599353" "titulo" => "Palabras clave" ] 4 => array:2 [ "identificador" => "sec0005" "titulo" => "Introduction" ] 5 => array:3 [ "identificador" => "sec0010" "titulo" => "Materials and methods" "secciones" => array:2 [ 0 => array:2 [ "identificador" => "sec0015" "titulo" => "Sample collection and taxonomic determination" ] 1 => array:2 [ "identificador" => "sec0020" "titulo" => "Morphometric analyses" ] ] ] 6 => array:3 [ "identificador" => "sec0025" "titulo" => "Description" "secciones" => array:2 [ 0 => array:2 [ "identificador" => "sec0030" "titulo" => "Type locality: Spring in Santiago Yosondúa, Oaxaca, Mexico" ] 1 => array:2 [ "identificador" => "sec0035" "titulo" => "Taxonomic summary" ] ] ] 7 => array:2 [ "identificador" => "sec0040" "titulo" => "Results" ] 8 => array:2 [ "identificador" => "sec0045" "titulo" => "Discussion" ] 9 => array:2 [ "identificador" => "xack196110" "titulo" => "Acknowledgements" ] 10 => array:1 [ "titulo" => "References" ] ] ] "pdfFichero" => "main.pdf" "tienePdf" => true "fechaRecibido" => "2014-08-13" "fechaAceptado" => "2015-07-22" "PalabrasClave" => array:2 [ "en" => array:1 [ 0 => array:4 [ "clase" => "keyword" "titulo" => "Keywords" "identificador" => "xpalclavsec599352" "palabras" => array:3 [ 0 => "New species" 1 => "Killifish" 2 => "Morphologic traits" ] ] ] "es" => array:1 [ 0 => array:4 [ "clase" => "keyword" "titulo" => "Palabras clave" "identificador" => "xpalclavsec599353" "palabras" => array:3 [ 0 => "Especie nueva" 1 => "Ciprinodóntido" 2 => "Caracteres morfológicos" ] ] ] ] "tieneResumen" => true "resumen" => array:2 [ "en" => array:2 [ "titulo" => "Abstract" "resumen" => "The family Profundulidae is one of the few families of freshwater fishes endemic to the Mesoamerican region. It includes the genus Profundulus that has been divided into two subgenera, Profundulus, with a wider distribution and Tlaloc, which presents a more limited range. The species currently recognized within the subgenus Profundulus are P. punctatus, P. oaxacae, P. guatemalensis and P. kreiseri. Results from a previous study using allozymes revealed a high degree of molecular differentiation of the populations inhabiting the Mixteca region (located at the north of Oaxaca in Mexico), compared to the rest of the species in the genus, especially regarding its geographically nearest neighbor (P. oaxacae). The main goal of this study was to evaluate the morphological distinctiveness and to assess the taxonomic identity of the populations from the Mixteca region in comparison with the morphological variation of P. oaxacae. Based on 22 measurements (17 morphometric and 5 meristic), we examined the morphological variation within and among species, including 50 individuals from 5 localities. Our results show a high degree of morphological differentiation between both populations in accordance to the molecular information reported before, supporting that populations from the Mixteca region are a new taxonomic entity, which is described in the present study." ] "es" => array:2 [ "titulo" => "Resumen" "resumen" => "La familia Profundulidae es una de las pocas familias de peces dulceacuícolas endémicas a la región de Mesoamérica. La familia incluye al género Profundulus, el cual a su vez incluye dos subgéneros: Profundulus, con una mayor distribución y Tlaloc, el cual presenta una distribución más restringida. Actualmente, dentro del género Profundulus se incluyen las siguientes especies: P. punctatus, P. oaxacae, P. guatemalensis y P. kreiseri. En un estudio previo basado en un análisis de aloenzimas para el subgénero Profundulus, se observó un alto grado de diferenciación entre las poblaciones de la región Mixteca, al norte del estado de Oaxaca en México, del resto de especies dentro del subgénero, particularmente de las poblaciones de la especie geográficamente más cercana (P. oaxacae). El objetivo principal del presente estudio es caracterizar la variación morfológica y evaluar la identidad taxonómica de las poblaciones de la región Mixteca comparándolas con la variación morfológica de P. oaxacae. Esta caracterización se llevó a cabo con base en 22 variables (17 morfométricas y 5 merísticas) en 50 individuos provenientes de 5 localidades. Nuestros resultados muestran que existe una clara diferenciación morfológica entre ambos grupos de poblaciones, en congruencia con evidencia molecular obtenida previamente, y que sustenta el reconocimiento de las poblaciones de la región de la Mixteca como una nueva especie, la cual es descrita en el presente estudio." ] ] "apendice" => array:1 [ 0 => array:1 [ "seccion" => array:1 [ 0 => array:4 [ "apendice" => "The following are the supplementary data to this article:<elsevierMultimedia ident="upi0005"></elsevierMultimedia>" "etiqueta" => "Appendix A" "titulo" => "Supplementary data" "identificador" => "sec0055" ] ] ] ] "multimedia" => array:9 [ 0 => array:7 [ "identificador" => "fig0005" "etiqueta" => "Figure 1" "tipo" => "MULTIMEDIAFIGURA" "mostrarFloat" => true "mostrarDisplay" => false "figura" => array:1 [ 0 => array:4 [ "imagen" => "gr1.jpeg" "Alto" => 1905 "Ancho" => 2666 "Tamanyo" => 561956 ] ] "descripcion" => array:1 [ "en" => "Map showing sampling points for the species: P. mixtlanensis sp. nov. (black circles) and P. oaxacae (white circles)." ] ] 1 => array:7 [ "identificador" => "fig0010" "etiqueta" => "Figure 2" "tipo" => "MULTIMEDIAFIGURA" "mostrarFloat" => true "mostrarDisplay" => false "figura" => array:1 [ 0 => array:4 [ "imagen" => "gr2.jpeg" "Alto" => 1893 "Ancho" => 1607 "Tamanyo" => 270651 ] ] "descripcion" => array:1 [ "en" => "Morphological comparison of the species of Profundulus: (a) Profundulus (P.) mixtlanensis sp. nov. and P. oaxacae individuals; (b) lateral view of the head of both species; (c) dorsal head view of both species; (d) holotype of P. mixtlanensis sp. nov." ] ] 2 => array:7 [ "identificador" => "fig0015" "etiqueta" => "Figure 3" "tipo" => "MULTIMEDIAFIGURA" "mostrarFloat" => true "mostrarDisplay" => false "figura" => array:1 [ 0 => array:4 [ "imagen" => "gr3.jpeg" "Alto" => 1177 "Ancho" => 1576 "Tamanyo" => 71934 ] ] "descripcion" => array:1 [ "en" => "Bivariate plot of PCA results from morphometric measurements after allometric regression. Solid circles correspond to P. mixtlanensis sp. nov. and white to P. oaxacae. PC1 explains 91.73% of the variance, whereas PC2 explains 2.79% of the variance." ] ] 3 => array:7 [ "identificador" => "fig0020" "etiqueta" => "Figure 4" "tipo" => "MULTIMEDIAFIGURA" "mostrarFloat" => true "mostrarDisplay" => false "figura" => array:1 [ 0 => array:4 [ "imagen" => "gr4.jpeg" "Alto" => 2486 "Ancho" => 2988 "Tamanyo" => 248813 ] ] "descripcion" => array:1 [ "en" => "Results from the discriminant function analysis (DFA): (a) proportions of correctly assigned individuals: 97% for P. mixtlanensis sp. nov. (individuals 1–35 in the graph) and 100% for P. oaxacae (individuals 36–50); (b) distribution frequencies of root factor from the DFA between species." ] ] 4 => array:7 [ "identificador" => "fig0025" "etiqueta" => "Figure 5" "tipo" => "MULTIMEDIAFIGURA" "mostrarFloat" => true "mostrarDisplay" => false "figura" => array:1 [ 0 => array:4 [ "imagen" => "gr5.jpeg" "Alto" => 3462 "Ancho" => 2883 "Tamanyo" => 315413 ] ] "descripcion" => array:1 [ "en" => "Frequency distribution of 5 meristic characters for P. mixtlanensis sp. nov. (black squares) and P. oaxacae (white squares)." ] ] 5 => array:7 [ "identificador" => "tbl0005" "etiqueta" => "Table 1" "tipo" => "MULTIMEDIATABLA" "mostrarFloat" => true "mostrarDisplay" => false "tabla" => array:1 [ "tablatextoimagen" => array:1 [ 0 => array:2 [ "tabla" => array:1 [ 0 => """ <table border="0" frame="\n \t\t\t\t\tvoid\n \t\t\t\t" class=""><thead title="thead"><tr title="table-row"><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">Map \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">Locality \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">N \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">Basin \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">Species proposed \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">Species by <a class="elsevierStyleCrossRefs" href="#bib0030">Miller (1955, 2005)</a> \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">UTM-X \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">UTM-Y \t\t\t\t\t\t\n \t\t\t\t</th></tr></thead><tbody title="tbody"><tr title="table-row"><td class="td" title="table-entry " align="char" valign="top">1 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Río Grande, Santiago Juxtlahuaca, Oaxaca, México \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">5 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Mixteco-Balsas \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. mixtlanensis \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. punctatus \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">605394 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">1910523 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry " align="char" valign="top">2 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Spring, Sola de Vega, Oaxaca. México. \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">– \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Atoyac \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. oaxacae \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. oaxacae \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">715433 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">1826989 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry " align="char" valign="top">3 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Río Cucharas, El Carmen Tuxtitlán, Putla de Guerrero, Oaxaca, México \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">7 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Atoyac \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. mixtlanensis \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. punctatus \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">615930 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">1884461 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry " align="char" valign="top">4 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Río Chiquito, Tlaxiaco, Oaxaca, México \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">15 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Mixteco-Balsas \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. oaxacae \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. oaxacae \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">643961 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">1905758 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry " align="char" valign="top">5 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Spring, Aldama, Oaxaca, México \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">15 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Atoyac \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. mixtlanensis \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. punctatus \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">648638 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">1875597 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry " align="char" valign="top">6 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Spring, Chalcatongo, Oaxaca, México \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">– \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Atoyac \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. mixtlanensis \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. punctatus \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">650543 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">1884349 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry " align="char" valign="top">7 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Spring, Santiago Yosondúa, Oaxaca. México \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">8 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Atoyac \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. mixtlanensis \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. punctatus \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">651751 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">1866154 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry " align="char" valign="top">8 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Río de las Grutas, San Sebastián de las Grutas, Oaxaca, México \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">– \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Atoyac \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. oaxacae \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">P. oaxacae \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">718060 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">1839603 \t\t\t\t\t\t\n \t\t\t\t</td></tr></tbody></table> """ ] "imagenFichero" => array:1 [ 0 => "xTab951137.png" ] ] ] ] "descripcion" => array:1 [ "en" => "Sampling localities, number of localities and number of individuals that were used in the morphologic analysis. Taxonomic designation is according to our study, and to <a class="elsevierStyleCrossRefs" href="#bib0030">Miller (1955, 2005)</a>." ] ] 6 => array:7 [ "identificador" => "tbl0010" "etiqueta" => "Table 2" "tipo" => "MULTIMEDIATABLA" "mostrarFloat" => true "mostrarDisplay" => false "tabla" => array:2 [ "leyenda" => "In bold highly significant p-values (p<0.005) and in cursives significant p-values (p<0.05 and p>0.005)." "tablatextoimagen" => array:1 [ 0 => array:2 [ "tabla" => array:1 [ 0 => """ <table border="0" frame="\n \t\t\t\t\tvoid\n \t\t\t\t" class=""><thead title="thead"><tr title="table-row"><th class="td-with-role" title="table-head ; entry_with_role_rowhead " align="left" valign="top" scope="col">Morphometric measurements \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " colspan="2" align="center" valign="top" scope="col" style="border-bottom: 2px solid black">Mean (standard error)</th><th class="td" title="table-head " colspan="4" align="center" valign="top" scope="col" style="border-bottom: 2px solid black">95% Confidence interval</th><th class="td" title="table-head " align="left" valign="top" scope="col">F \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col">p-Value \t\t\t\t\t\t\n \t\t\t\t</th></tr><tr title="table-row"><th class="td" title="table-head " align="" valign="top" scope="col"> \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="" valign="top" scope="col"> \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="" valign="top" scope="col"> \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " colspan="2" align="center" valign="top" scope="col" style="border-bottom: 2px solid black">Upper limit</th><th class="td" title="table-head " colspan="2" align="center" valign="top" scope="col" style="border-bottom: 2px solid black">Lower limit</th><th class="td" title="table-head " align="" valign="top" scope="col"> \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="" valign="top" scope="col"> \t\t\t\t\t\t\n \t\t\t\t</th></tr><tr title="table-row"><th class="td" title="table-head " align="" valign="top" scope="col" style="border-bottom: 2px solid black"> \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">P. mixtlanensis \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">P. oaxacae \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">P. mixtlanensis \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">P. oaxacae \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">P. mixtlanensis \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">P. oaxacae \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="" valign="top" scope="col" style="border-bottom: 2px solid black"> \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="" valign="top" scope="col" style="border-bottom: 2px solid black"> \t\t\t\t\t\t\n \t\t\t\t</th></tr></thead><tbody title="tbody"><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Standard length (SL) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">63.48 (1.70) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">47.18 (2.41) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">66.91 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">52.04 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">60.05 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">42.33 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">30.5 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Head length (HL) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">17.51 (0.52) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">13.72 (0.74) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">18.57 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">15.22 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">16.46 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">12.22 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">17.34 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pre-doral distance (PrDD) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">42.11 (1.11) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">31.96 (1.58) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">44.36 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">35.15 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">39.87 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">28.78 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">27.53 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pre-anal distance (PrPA) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">42.03 (1.09) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">32.60 (1.54) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">44.23 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">35.72 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">39.83 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">29.49 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">24.74 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Eye diameter (ED) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">4.15 (0.07) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">3.41 (0.11) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">4.31 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">3.64 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">3.99 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">3.18 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">28.63 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pre-orbital length (PrOL) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">4.80 (0.16) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">3.36 (0.23) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">5.13 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">3.83 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">4.47 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">2.89 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">25.35 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pre-pectoral distance (PrPP) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">17.77 (0.53) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">14.08 (0.75) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">18.84 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">15.60 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">16.70 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">12.56 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">15.98 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pre-ventral distance (PrPV) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">32.81 (0.93) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">25.06 (1.32) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">34.69 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">27.73 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">30.93 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">22.39 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">22.88 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Body depth (BD) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">17.46 (0.72) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">14.70 (1.02) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">18.91 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">16.76 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">16.00 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">12.64 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">4.85 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.033 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Dorsal fin length (DFL) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">10.07 (0.32) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">7.67 (0.46) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">10.74 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">8.60 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">9.41 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">6.73 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">17.86 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pectoral-ventral length (PVL) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">16.74 (0.63) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">12.54 (0.90) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">18.02 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">14.37 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">15.45 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">10.72 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">14.28 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Anal fin length (AFL) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">10.33 (0.32) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">6.95 (0.46) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">11 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">7.89 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">9.67 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">6.02 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">35.30 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pectoral fin length (PFL) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">9.76 (0.23) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">7.19 (0.33) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">10.23 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">7.86 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">9.29 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">6.52 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">39.77 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Ventral–anal length (VAL) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">9.59 (0.27) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">7.75 (0.38) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">10.13 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">8.52 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">9.04 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">6.98 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">15.34 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Body least depth (BLD) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">8.85 (0.28) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">7.407 (0.40) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">9.42 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">8.21 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">8.28 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">6.59 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">8.61 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.005 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Anal peduncle length (APL) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">10.53 (0.35) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">7.024 (0.50) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">11.25 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">8.04 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">9.81 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">6.00 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">32.22 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Caudal peduncle length (CPL) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">10.89 (0.33) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">8.007 (0.46) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">11.55 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">8.94 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">10.23 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">7.06 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">25.47 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry " colspan="9" align="left" valign="top"></td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Meristic counts \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="" valign="top"> \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="" valign="top"> \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " colspan="2" align="center" valign="top">Max</td><td class="td" title="table-entry " colspan="2" align="center" valign="top">Min</td><td class="td" title="table-entry " align="" valign="top"> \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="left" valign="top">Mann–Whitney U test \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Number of lateral scales (E) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">33.31 (0.20) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">30.93(0.283) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">35 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">32 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">30 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">29 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="" valign="top"> \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Doral fin rays (D) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">11.45 (0.12) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">11.4 (0.130) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">13 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">12 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">10 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">11 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="" valign="top"> \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.725 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Anal fin rays (A) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">14.45 (0.27) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">13.13 (0.215) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">21 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">14 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">12 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">12 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="" valign="top"> \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.001 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pectoral fin rays (P) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">15.94 (0.17) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">14.33 (0.159) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">17 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">16 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">14 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">14 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="" valign="top"> \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Ventral fin rays (V) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">6.05 (0.03) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">5.93 (0.066) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">7 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">6 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">6 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">5 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="" valign="top"> \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.105 \t\t\t\t\t\t\n \t\t\t\t</td></tr></tbody></table> """ ] "imagenFichero" => array:1 [ 0 => "xTab951138.png" ] ] ] ] "descripcion" => array:1 [ "en" => "Mean, standard error and range values for morphometric measurements (mm) and meristic counts of P. mixtlanensis and P. oaxacae. p-Values from the ANOVA test for morphometric measurements and those from the Mann–Whitney U test for meristic counts are provided." ] ] 7 => array:7 [ "identificador" => "tbl0015" "etiqueta" => "Table 3" "tipo" => "MULTIMEDIATABLA" "mostrarFloat" => true "mostrarDisplay" => false "tabla" => array:2 [ "leyenda" => "SL, standard length; HL, head length; PrPA, pre-anal distance; PrPP, pre-pectoral distance; BD, body depth; AFL, anal fin length; VAL, ventral–anal length; BLD, body least depth; APL, anal peduncle length; PrOL, pre-orbital length, ED, eye diameter. In bold highly significant p-values (p<0.005) and in cursives significant p-values (p<0.05 and p>0.005)." "tablatextoimagen" => array:1 [ 0 => array:2 [ "tabla" => array:1 [ 0 => """ <table border="0" frame="\n \t\t\t\t\tvoid\n \t\t\t\t" class=""><thead title="thead"><tr title="table-row"><th class="td-with-role" title="table-head ; entry_with_role_rowhead " align="left" valign="top" scope="col">Proportion \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col">N \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " colspan="2" align="center" valign="top" scope="col" style="border-bottom: 2px solid black">Mean (SE)</th><th class="td" title="table-head " colspan="2" align="center" valign="top" scope="col" style="border-bottom: 2px solid black">Upper limit</th><th class="td" title="table-head " colspan="2" align="center" valign="top" scope="col" style="border-bottom: 2px solid black">Lower limit</th><th class="td" title="table-head " align="left" valign="top" scope="col">p \t\t\t\t\t\t\n \t\t\t\t</th></tr><tr title="table-row"><th class="td" title="table-head " align="" valign="top" scope="col" style="border-bottom: 2px solid black"> \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="" valign="top" scope="col" style="border-bottom: 2px solid black"> \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">P. mixtlanensis \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">P. oaxacae \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">P. mixtlanensis \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">P. oaxacae \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">P. mixtlanensis \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">P. oaxacae \t\t\t\t\t\t\n \t\t\t\t</th><th class="td" title="table-head " align="" valign="top" scope="col" style="border-bottom: 2px solid black"> \t\t\t\t\t\t\n \t\t\t\t</th></tr></thead><tbody title="tbody"><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">PrPA/SL \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">50 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.66 (0.0025) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.687 (0.0050) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.667 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.698 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.657 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.676 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">PrPP/SL \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">50 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.28 (0.0040) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.302 (0.00376) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.289 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.310 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.272 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.294 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">BD/SL \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">50 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.27 (0.006) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.310 (0.0043) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.285 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.320 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.260 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.301 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">AFL/SL \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">50 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.16 (0.0025) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.151 (0.0039) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.169 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.159 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.159 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.142 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.011 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">VAL/SL \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">50 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.15 (0.0021) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.163 (0.0048) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.155 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.173 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.146 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.153 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.041 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">BLD/SL \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">50 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.14 (0.0015) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.157 (0.0015) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.143 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.161 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.136 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.154 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.000 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">APL/SL \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">50 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.16 (0.0024) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.148 (0.0032) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.166 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.155 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.156 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.142 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.001 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">HL/SL \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">50 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.27 (0.0044) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.293 (0.0029) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.284 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.299 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.266 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.286 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.002 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">PrOL/HL \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">50 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.28 (0.0103) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.244 (0.0071) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.305 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.259 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.263 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.228 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.007 \t\t\t\t\t\t\n \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">ED/SL \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">50 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.27 (0.0044) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.293 (0.0029) \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.284 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.299 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.266 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.286 \t\t\t\t\t\t\n \t\t\t\t</td><td class="td" title="table-entry " align="char" valign="top">0.002 \t\t\t\t\t\t\n \t\t\t\t</td></tr></tbody></table> """ ] "imagenFichero" => array:1 [ 0 => "xTab951136.png" ] ] ] ] "descripcion" => array:1 [ "en" => "Mean, standard error and range values for proportions of morphometric variables between P. oaxacae and P. mixtlanensis. p-Values correspond to the Mann–Whitney U test." ] ] 8 => array:5 [ "identificador" => "upi0005" "tipo" => "MULTIMEDIAECOMPONENTE" "mostrarFloat" => false "mostrarDisplay" => true "Ecomponente" => array:2 [ "fichero" => "mmc1.docx" "ficheroTamanyo" => 34697 ] ] ] "bibliografia" => array:2 [ "titulo" => "References" "seccion" => array:1 [ 0 => array:2 [ "identificador" => "bibs0005" "bibliografiaReferencia" => array:11 [ 0 => array:3 [ "identificador" => "bib0005" "etiqueta" => "Álvarez del Villar, 1970" "referencia" => array:1 [ 0 => array:2 [ "contribucion" => array:1 [ 0 => array:2 [ "titulo" => "Peces mexicanos (claves)" "autores" => array:1 [ 0 => array:2 [ "etal" => false "autores" => array:1 [ 0 => "J. 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Doadrio" ] ] ] ] ] "host" => array:1 [ 0 => array:2 [ "doi" => "10.1016/j.ympev.2015.09.002" "Revista" => array:7 [ "tituloSerie" => "Molecular Phylogenetics and Evolution" "fecha" => "2015" "volumen" => "94" "numero" => "Part A" "paginaInicial" => "242" "paginaFinal" => "251" "link" => array:1 [ 0 => array:2 [ "url" => "https://www.ncbi.nlm.nih.gov/pubmed/26364972" "web" => "Medline" ] ] ] ] ] ] ] ] 9 => array:3 [ "identificador" => "bib0050" "etiqueta" => "Ruiz-Campos et al., 2003" "referencia" => array:1 [ 0 => array:2 [ "contribucion" => array:1 [ 0 => array:2 [ "titulo" => "Morphometric variation of wild trout populations from northwestern Mexico (Pisces: Salmonidae)" "autores" => array:1 [ 0 => array:2 [ "etal" => false "autores" => array:5 [ 0 => "G. Ruiz-Campos" 1 => "F. Camarena-Rosales" 2 => "A. Varela-Romero" 3 => "S. Sánchez-Gonzáles" 4 => "J. 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Taxonomy and systematics

Una especie nueva de ciprinodontiforme de la familia Profundulidae de las tierras altas de la región Mixteca, México

Claudia Patricia Ornelas-Garcíaa,b,

Corresponding author

patriciaornelasg@gmail.com

Corresponding author.

, Emilio Martínez-Ramírezc, Ignacio Doadriod

a Departamento de Zoología, Facultad de Ciencias Naturales, Universidad Autónoma de Querétaro, Av. de las Ciencias s/n, 76230 Juriquilla, Querétaro, Mexico

b Departamento de Zoología, Universidad Nacional Autónoma de México, Trecer Circuito s/n, Ciudad Universitaria, Coyoacán, 04510 Mexico City, Mexico

c Departamento de Investigación, Centro Interdisciplinario de Investigación para el Desarrollo Integral Regional Unidad Oaxaca, Instituto Politécnico Nacional, Calle Hornos N° 1003, Municipio Santa Cruz Xoxocotlán, 71230 Oaxaca, Mexico

d Departamento de Biodiversidad y Biología Evolutiva, Museo Nacional de Ciencias Naturales, CSIC, José Gutiérrez Abascal 2, 28006 Madrid, Spain

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    "textoCompleto" => "<span class="elsevierStyleSections"><span id="sec0005" class="elsevierStyleSection elsevierViewall"><span class="elsevierStyleSectionTitle" id="sect0025">Introduction</span><p id="par0005" class="elsevierStylePara elsevierViewall">The family Profundulidae is one of few families of freshwater fishes endemic to the Mesoamerican region&#46; It has been suggested that its history goes back to the Pliocene&#44; or even the Miocene &#40;<a class="elsevierStyleCrossRef" href="#bib0030">Miller&#44; 1955</a>&#41;&#46; Currently&#44; most species in the family exhibit low abundances&#44; and their distribution is restricted to the headwaters and spring-fed-tributaries of rivers in southern Mexico&#44; El Salvador&#44; Honduras and Guatemala&#44; from both Pacific and Atlantic versants&#46; Species diversity in the family is fairly low&#44; consisting of 8 species in 2 genera&#58; <span class="elsevierStyleItalic">Profundulus</span> and <span class="elsevierStyleItalic">Tlaloc</span> &#40;<a class="elsevierStyleCrossRef" href="#bib0045">Morcillo&#44; Ornelas-Garc&#237;a&#44; Alcaraz&#44; Matamoros&#44; &#38; Doadrio&#44; 2015</a>&#41;&#46; In the genus <span class="elsevierStyleItalic">Tlaloc</span>&#44; 4 species are currently recognized&#58; <span class="elsevierStyleItalic">T&#46; candalarius</span>&#44; <span class="elsevierStyleItalic">T</span>&#46; <span class="elsevierStyleItalic">hildebrandi</span>&#44; <span class="elsevierStyleItalic">T&#46; labialis</span> and <span class="elsevierStyleItalic">T&#46; portillorum</span>&#44; all of which exhibit limited distributions&#46; The genus <span class="elsevierStyleItalic">Profundulus</span> also has 4 species&#44; namely <span class="elsevierStyleItalic">P&#46; punctatus</span>&#44; <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#44; <span class="elsevierStyleItalic">P&#46; guatemalensis</span> and <span class="elsevierStyleItalic">P&#46; kreiseri</span>&#44; but has a decidedly wider distribution range&#46;</p><p id="par0010" class="elsevierStylePara elsevierViewall">In an early study dealing with systematics of Profundulidae&#44; <a class="elsevierStyleCrossRef" href="#bib0030">Miller &#40;1955&#41;</a> pointed out a wide degree of morphological variation&#44; being more evident within the <span class="elsevierStyleItalic">P&#46; punctatus</span> &#8220;group&#8221;&#44; in which more than one species might be recognized&#46; Years later&#44; within the <span class="elsevierStyleItalic">P&#46; punctatus</span> &#8220;group&#8221; <a class="elsevierStyleCrossRef" href="#bib0035">Miller &#40;2005&#41;</a> corroborated the validity of <span class="elsevierStyleItalic">P&#46; oaxacae</span> &#40;<a class="elsevierStyleCrossRef" href="#bib0025">Meek&#44; 1902</a>&#41;&#44; whose distribution was restricted to altitudes above 1&#44;550<span class="elsevierStyleHsp" style=""></span>m in the R&#237;o Atoyac &#8211; Verde Basin&#44; Oaxaca&#44; in southern Mexico &#40;<a class="elsevierStyleCrossRef" href="#bib0020">Mart&#237;nez-Ram&#237;rez&#44; Doadrio-Villarejo&#44; &#38; de Sostoa-Fern&#225;ndez&#44; 2004</a>&#41;&#46; Furthermore&#44; in a genetic population analysis of the subgenus <span class="elsevierStyleItalic">Profundulus</span> based on allozymes &#40;<a class="elsevierStyleCrossRef" href="#bib0010">Doadrio&#44; Carmona&#44; Mart&#237;nez&#44; &#38; De Sostoa&#44; 1999</a>&#41;&#44; at least 5 different lineages were recovered&#46; Of these&#44; 2 distinct lineages were found in the Mixteca region and were well differentiated from <span class="elsevierStyleItalic">P&#46; punctatus</span>&#46; One of them corresponded to <span class="elsevierStyleItalic">P&#46; oaxacae</span> and the other one included a clade named <span class="elsevierStyleItalic">P&#46;</span> sp&#46; nov&#46; 1&#44; which corresponded to populations from Mixteca headwaters &#40;with an elevation over 700<span class="elsevierStyleHsp" style=""></span>m&#41; and showed a high degree of molecular divergence from the rest of species of the subgenus <span class="elsevierStyleItalic">Profundulus</span> &#40;Nei&#39;s distances <span class="elsevierStyleItalic">D</span><span class="elsevierStyleHsp" style=""></span>&#61;<span class="elsevierStyleHsp" style=""></span>0&#46;165<span class="elsevierStyleHsp" style=""></span>&#8722;<span class="elsevierStyleHsp" style=""></span>0&#46;845&#41;&#46; In fact&#44; based on the aforementioned analysis&#44; <span class="elsevierStyleItalic">P&#46;</span> sp&#46; nov&#46; 1 was recovered as a sister clade to the rest of species in the <span class="elsevierStyleItalic">P&#46; punctatus</span> &#8220;group&#8221;&#44; where <span class="elsevierStyleItalic">P&#46; oaxacae</span> is included&#46;</p><p id="par0015" class="elsevierStylePara elsevierViewall">As a consequence of the striking morphological differences between <span class="elsevierStyleItalic">P&#46; oaxacae</span> and <span class="elsevierStyleItalic">P&#46; punctatus</span>&#44; Miller recognized the former as a valid species &#40;<a class="elsevierStyleCrossRef" href="#bib0035">Miller&#44; 2005</a>&#41;&#46; These differences include the number of scales around the body &#40;25&#8211;28 in <span class="elsevierStyleItalic">P&#46; punctatus vs</span>&#46; 30 in <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#41;&#44; and the lateral color pattern &#40;prominent round spots in <span class="elsevierStyleItalic">P&#46; punctatus vs</span>&#46; narrow weak and vertical marks in <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#41;&#46;</p><p id="par0020" class="elsevierStylePara elsevierViewall">Another aspect clearly separating this pair of species is their distribution&#44; in which <span class="elsevierStyleItalic">P&#46; oaxacae</span> is restricted to the upper part of the r&#237;o Verde basin&#44; Oaxaca while <span class="elsevierStyleItalic">P&#46; punctatus</span> inhabits the lower part of the same basin&#46; <span class="elsevierStyleItalic">P&#46;</span> sp&#46; nov&#46; 1 also inhabits the upper part of the r&#237;o Verde basin&#44; but shows a high genetic distinctiveness regarding both&#44; <span class="elsevierStyleItalic">P&#46; oaxacae</span> and <span class="elsevierStyleItalic">P&#46; punctatus</span> &#40;<a class="elsevierStyleCrossRef" href="#bib0010">Doadrio et al&#46;&#44; 1999</a>&#41;&#46; Consequently&#44; in this study we evaluated the morphological distinctiveness between <span class="elsevierStyleItalic">P&#46; oaxacae</span> and <span class="elsevierStyleItalic">P&#46;</span> sp&#46; nov&#46; 1&#44; and determined whether morphological variation corresponds to the genetic differentiation previously documented by <a class="elsevierStyleCrossRef" href="#bib0010">Doadrio et al&#46; &#40;1999&#41;</a>&#46;</p></span><span id="sec0010" class="elsevierStyleSection elsevierViewall"><span class="elsevierStyleSectionTitle" id="sect0030">Materials and methods</span><span id="sec0015" class="elsevierStyleSection elsevierViewall"><span class="elsevierStyleSectionTitle" id="sect0035">Sample collection and taxonomic determination</span><p id="par0025" class="elsevierStylePara elsevierViewall">A total of 5 localities within the Mixteca region were surveyed using cast nets and electrofishing &#40;<a class="elsevierStyleCrossRef" href="#fig0005">Fig&#46; 1</a>&#41;&#46; A total of 50 specimens were collected&#44; with a range of individuals per locality from 5 to 15 &#40;<a class="elsevierStyleCrossRef" href="#tbl0005">Table 1</a>&#41;&#46; Fin clips were obtained and individuals tagged&#46; Fin clips and voucher specimens were preserved in 95&#37; ethanol and deposited in Colecci&#243;n Nacional de Peces&#44; Instituto de Biolog&#237;a&#44; Universidad Nacional Aut&#243;noma de M&#233;xico &#40;CNPE-IBUNAM&#41;&#44; Colecci&#243;n de Peces&#44; El Colegio de la Frontera Sur &#40;ECOSC&#41;&#44; Museo Nacional de Ciencias Naturales de Madrid&#44; Espa&#241;a &#40;MNCN&#41; and Colecci&#243;n de Peces del Centro de M&#233;xico Dr&#46; Edmundo D&#237;az Pardo&#44; Universidad Aut&#243;noma de Quer&#233;taro&#44; Mexico &#40;CPCMEDP-UAQ&#41;&#46; Localities for <span class="elsevierStyleItalic">P&#46;</span> sp&#46; nov&#46; 1 were determined based on a previous molecular study &#40;<a class="elsevierStyleCrossRef" href="#tbl0005">Table 1</a>&#59; <a class="elsevierStyleCrossRef" href="#bib0010">Doadrio et al&#46;&#44; 1999</a>&#41;&#46; For <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#44; we measured 15 specimens from Rio Chiquito&#44; Tlaxiaco&#44; in the Mixteco-Balsas basin&#46; <span class="elsevierStyleItalic">P&#46; oaxacae</span> were identified using Miller&#39;s keys &#40;<a class="elsevierStyleCrossRef" href="#bib0035">Miller&#44; 2005</a>&#41;&#46;</p><elsevierMultimedia ident="fig0005"></elsevierMultimedia><elsevierMultimedia ident="tbl0005"></elsevierMultimedia></span><span id="sec0020" class="elsevierStyleSection elsevierViewall"><span class="elsevierStyleSectionTitle" id="sect0040">Morphometric analyses</span><p id="par0030" class="elsevierStylePara elsevierViewall">For comparison between species groups&#44; we performed 17 morphometric measurements and 5 meristic counts in 50 individuals from 5 localities &#40;<a class="elsevierStyleCrossRef" href="#fig0005">Fig&#46; 1</a>&#44; <a class="elsevierStyleCrossRefs" href="#tbl0005">Tables 1 and 2</a>&#41;&#46; In order to identify the variables that contributed the most to the variation among populations&#44; we performed principal component analysis &#40;PCA&#41; on the morphometric dataset&#44; based on a covariance matrix&#46; Finally&#44; we performed discriminant function analysis &#40;DFA&#41; and linear discriminant analysis &#40;LDA&#41; in order to evaluate each specimen&#44; assign it to a species and assess the degree of separation between the groups&#46;</p><elsevierMultimedia ident="tbl0010"></elsevierMultimedia><p id="par0035" class="elsevierStylePara elsevierViewall">For both analyses&#44; measurements were standardized using the general allometric equation&#58; Ms<span class="elsevierStyleHsp" style=""></span>&#61;<span class="elsevierStyleHsp" style=""></span>Mo &#40;Ls&#47;Lo&#41;<span class="elsevierStyleSup"><span class="elsevierStyleItalic">b</span></span>&#44; where Ms<span class="elsevierStyleHsp" style=""></span>&#61;<span class="elsevierStyleHsp" style=""></span>standardized measurement&#44; Mo<span class="elsevierStyleHsp" style=""></span>&#61;<span class="elsevierStyleHsp" style=""></span>measured character length &#40;mm&#41;&#44; Ls<span class="elsevierStyleHsp" style=""></span>&#61;<span class="elsevierStyleHsp" style=""></span>overall &#40;arithmetic&#41; mean of the standard component &#40;standard length &#8211; SL or head length &#8211; HL&#41; for all individuals of each taxon&#44; Lo<span class="elsevierStyleHsp" style=""></span>&#61;<span class="elsevierStyleHsp" style=""></span>standard component of the specimen&#46; For each character &#8220;<span class="elsevierStyleItalic">b</span>&#8221; was estimated using the non-linear equation&#44; <span class="elsevierStyleItalic">M</span><span class="elsevierStyleHsp" style=""></span>&#61;<span class="elsevierStyleHsp" style=""></span><span class="elsevierStyleItalic">aLb</span> as the slope of the regression of log<span class="elsevierStyleHsp" style=""></span>Mo on log<span class="elsevierStyleHsp" style=""></span>Lo &#40;<a class="elsevierStyleCrossRefs" href="#bib0015">Elliot&#44; Haskard&#44; &#38; Koslow&#44; 1995&#59; Ruiz-Campos&#44; Camarena-Rosales&#44; Varela-Romero&#44; S&#225;nchez-Gonz&#225;les&#44; &#38; De La Rosa-V&#233;lezv&#44; 2003</a>&#41; &#40;logistic regression in Supplementary material 1&#41;&#46; Abbreviations for morphological measurements are given in <a class="elsevierStyleCrossRef" href="#tbl0010">Table 2</a>&#46;</p><p id="par0040" class="elsevierStylePara elsevierViewall">Two-way analysis of variance &#40;ANOVA&#41; was carried out considering sex and species as factors to evaluate the influence of sexual dimorphism&#44; and the sex<span class="elsevierStyleHsp" style=""></span>&#215;<span class="elsevierStyleHsp" style=""></span>species interaction was included to test if dimorphism varied significantly between species groups for the 17 morphometric measurements&#46; Proportions between measurements were calculated in relation to SL and HL &#40;<a class="elsevierStyleCrossRef" href="#tbl0015">Table 3</a>&#41;&#46; Frequencies of the 5 meristic variables were compared between species groups&#46; Mann&#8211;Whitney <span class="elsevierStyleItalic">U</span> tests were carried out on the 5 meristic variables as well as on the morphometric proportion measurements&#46; Statistica v&#46;6&#46;0 &#40;<a class="elsevierStyleCrossRef" href="#bib0055">Statistica&#44; 2001</a>&#41; was used for all statistical analyses&#46;</p><elsevierMultimedia ident="tbl0015"></elsevierMultimedia><p id="par0045" class="elsevierStylePara elsevierViewall">Diagnostic measurements were expressed as percent of the standard length &#40;SL&#41; &#40;<a class="elsevierStyleCrossRef" href="#bib0005">&#193;lvarez del Villar&#44; 1970</a>&#41;&#46; Additionally&#44; the diagnosis was completed with other relevant features regarding morphology &#40;<span class="elsevierStyleItalic">i&#46;e&#46;</span>&#44; body profile&#44; fin shapes&#44; among others&#41;&#44; pigmentation and ecology&#46;</p></span></span><span id="sec0025" class="elsevierStyleSection elsevierViewall"><span class="elsevierStyleSectionTitle" id="sect0045">Description</span><p id="par0050" class="elsevierStylePara elsevierViewall">Profundulus mixtlanesis <span class="elsevierStyleItalic">Ornelas-Garc&#237;a&#44; Mart&#237;nez &#38; Doadrio&#44; new species &#40;</span><a class="elsevierStyleCrossRef" href="#fig0010"><span class="elsevierStyleItalic">Fig&#46; 2</span></a><span class="elsevierStyleItalic">&#41;&#46;</span></p><elsevierMultimedia ident="fig0010"></elsevierMultimedia><span id="sec0030" class="elsevierStyleSection elsevierViewall"><span class="elsevierStyleSectionTitle" id="sect0050">Type locality&#58; Spring in Santiago Yosond&#250;a&#44; Oaxaca&#44; Mexico</span><p id="par0055" class="elsevierStylePara elsevierViewall">As a member of the <span class="elsevierStyleItalic">Profundulus</span> subgenus this species shows a preorbital region nearly covered with well-developed scales&#44; a humeral spot&#44; and a densely scaled basal half of the caudal fin&#46; <span class="elsevierStyleItalic">P&#46; mixtlanensis</span> sp&#46; nov&#46; differs from other species in the genus by having a pronounced lower jaw&#44; which markedly differentiates it from <span class="elsevierStyleItalic">P&#46; oaxacae</span> and the remaining species within the <span class="elsevierStyleItalic">P&#46; punctatus</span> &#8220;group&#8221; <span class="elsevierStyleItalic">sensu</span><a class="elsevierStyleCrossRef" href="#bib0030">Miller &#40;1955&#41;</a>&#46; Another striking difference between the new species and other species in the subgenus&#44; especially <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#44; is the absence of dark body spotting on the posterior half of the body caused by melanization of the base of the scales&#46; In lateral view&#44; the dorsal profile of the head shows a concavity just above the supra-occipital crest&#44; which quickly reaches a flat profile posteriorly&#44; an attribute that becomes more pronounced with age&#46; In general terms&#44; <span class="elsevierStyleItalic">P&#46; mixtlanesis</span> depicts a longer body and has a lower height than the other species in the subgenus &#40;see <a class="elsevierStyleCrossRef" href="#tbl0015">Table 3</a>&#41;&#46; Body height is &#60;2&#47;3 of SL&#44; and pre-pectoral distance &#60;1&#47;3 of SL&#44; in contrast the <span class="elsevierStyleItalic">P&#46; oaxacae</span> proportions are significantly higher reflecting larger body height and pre-pectoral distance for the <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#46; The distance between ventral and anal fins is 15&#37; of SL&#44; lower than that of <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#46; Caudal peduncle depth is less than in <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#44; being 13&#37; of SL &#40;15&#37; of SL in <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#41;&#46; The postanal length is 15&#37; of SL&#44; larger than the rest of species &#40;data not shown&#41;&#46; The head is shorter than in <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#44; 26&#8211;28&#37; of SL in contrast to 28&#8211;30&#37; of SL in <span class="elsevierStyleItalic">P&#46; mixtlanensis</span>&#46; Preorbital length&#44; 26&#8211;30&#37; of HL&#44; is longer than in <span class="elsevierStyleItalic">P&#46; oaxacae&#46;</span> Eye diameter is smaller than in <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#44; 26&#8211;28&#37; of SL&#46; The anal fin is longer than in <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#44; 16&#8211;17&#37; of SL&#46; Among meristic characteristics&#44; the following are diagnostic&#58; 30&#8211;35 lateral scales&#44; 12&#8211;21 rays in the anal fin&#44; and 14&#8211;17 rays in the pectoral fin&#46;</p></span><span id="sec0035" class="elsevierStyleSection elsevierViewall"><span class="elsevierStyleSectionTitle" id="sect0055">Taxonomic summary</span><p id="par0060" class="elsevierStylePara elsevierViewall"><span class="elsevierStyleItalic">Disposition of types</span>&#46; Holotype CNPE-IBUNAM 19112 &#40;Female&#44; T1339&#41; and allotype CNPE-IBUNAM 19113 &#40;male&#44; T1340&#41;&#44; Spring system in Santiago Yosond&#250;a&#44; Oaxaca&#44; M&#233;xico&#46; The system is composed of five small springs&#44; &#8764;1<span class="elsevierStyleHsp" style=""></span>m in diameter and less than 1<span class="elsevierStyleHsp" style=""></span>m deep each&#46; Deposited at the CNPE-IBUNAM&#46;</p><p id="par0065" class="elsevierStylePara elsevierViewall"><span class="elsevierStyleItalic">Paratypes</span>&#46; &#40;45&#41;&#44; Spring&#44; Aldama&#44; Oaxaca &#40;15&#44; CPCMEDP-UAQ&#58;001&#47;T1317-31&#41;&#59; r&#237;o Cucharas&#44; El Carmen Tuxtitl&#225;n&#44; Putla de Guerrero&#44; Oaxaca &#40;3&#44; CPCMEDP-UAQ&#58;002&#47;1332-1335&#41;&#59; Spring&#44; Santiago Yosond&#250;a&#44; Oaxaca &#40;6&#44; 3 CNPE-IBUNAM 19114&#44; 3&#59; CPCMEDP-UAQ&#58;003&#47;T1340-1346-&#41;&#59; r&#237;o Grande&#44; Santiago Juxtlahuaca&#44; Oaxaca &#40;3&#44; ECOSC 7562-1&#47;ECOSC 7562-3 and 2&#44; CPCMEDP-UAQ&#58;004&#47;T1350-51&#44; CPCMEDP-UAQ&#41;&#46; R&#237;o Cucharas &#40;10&#59; MNCN&#58;131746-53&#41;&#46;</p><p id="par0070" class="elsevierStylePara elsevierViewall"><span class="elsevierStyleItalic">Distribution</span>&#46; Springs of the Pacific slope of the Mixteca region of Oaxaca&#44; Mexico in the following river systems&#58; Colorado River&#44; Atoyaquillo River&#44; the Atoyac river basin&#44; and the Mixteco river basin &#40;<a class="elsevierStyleCrossRef" href="#fig0005">Fig&#46; 1</a>&#41;&#46; Most of the populations for the species were found above 2&#44;000<span class="elsevierStyleHsp" style=""></span>m&#44; however&#44; they occur at altitudes between 710 and 2&#44;300<span class="elsevierStyleHsp" style=""></span>m&#46;</p><p id="par0075" class="elsevierStylePara elsevierViewall"><span class="elsevierStyleItalic">Etymology</span>&#46; The epithet &#8220;mixtlanensis&#8221; derives from the word &#8220;Mixtlan&#8221; in the Nahuatl language&#46; The Tenoch culture used to call Mixtlan the region where the new species is found&#46; Mixtlan is a compound word&#44; which is made up of the words Mixtli &#40;cloud&#41; and -Tlan &#40;place&#41;&#46; Hence it means &#8220;the place of the clouds&#8221; &#40;<a class="elsevierStyleCrossRef" href="#bib0040">Montemayor&#44; 2008</a>&#41;&#46;</p></span></span><span id="sec0040" class="elsevierStyleSection elsevierViewall"><span class="elsevierStyleSectionTitle" id="sect0060">Results</span><p id="par0080" class="elsevierStylePara elsevierViewall">Morphometric analyses showed a separation between the two species groups tested&#46; The two-way ANOVA using 17 morphometric measurements showed significant differences between species for most of the comparisons &#40;<span class="elsevierStyleItalic">p</span><span class="elsevierStyleHsp" style=""></span>&#8804;<span class="elsevierStyleHsp" style=""></span>0&#46;001&#44; <a class="elsevierStyleCrossRef" href="#tbl0010">Table 2</a>&#41;&#46; Analyses however&#44; did not disclose any significant differences in sex &#40;<span class="elsevierStyleItalic">p</span><span class="elsevierStyleHsp" style=""></span>&#62;<span class="elsevierStyleHsp" style=""></span>0&#46;05&#41;&#44; or sex<span class="elsevierStyleHsp" style=""></span>&#215;<span class="elsevierStyleHsp" style=""></span>species interaction &#40;<span class="elsevierStyleItalic">p</span><span class="elsevierStyleHsp" style=""></span>&#62;<span class="elsevierStyleHsp" style=""></span>0&#46;05&#44; data not shown&#41;&#46; Because results did not support the existence of sexual dimorphism within or between species&#44; all posterior analyses were performed without sex separations&#46; Ordination using PCA allowed for identification of two separate clusters of points&#46; Each cluster corresponded to <span class="elsevierStyleItalic">P&#46; oaxacae</span> and to <span class="elsevierStyleItalic">P&#46;</span> sp&#46; nov&#46; 1 &#40;<a class="elsevierStyleCrossRef" href="#fig0015">Fig&#46; 3</a>&#41;&#46; PCI explained 91&#46;73&#37; of the variance&#44; where the most important variables in the formation of the groups were SL&#44; PrOL and ED in the positive side of PCI&#44; and PrPD&#44; PrPA and PrPV in the negative side&#46; PCII explained 2&#46;79&#37; of the variance&#44; in which the most important variables were PrPD&#44; PrPA&#44; PrPP and BD&#44; whereas PCIII explained 1&#46;70&#37; of the variance&#44; with the most important variables PrPP&#44; HL&#44; PVL and VAL &#40;Eigenvalues in Supplementary material 4&#41;&#46;</p><elsevierMultimedia ident="fig0015"></elsevierMultimedia><p id="par0085" class="elsevierStylePara elsevierViewall">DFA also showed significant discrimination between species&#46; All individuals assigned to <span class="elsevierStyleItalic">P&#46; oaxacae</span> were correctly classified whereas assignation to <span class="elsevierStyleItalic">P&#46;</span> sp&#46; nov&#46; 1 was 97&#46;14&#37; correct &#40;<a class="elsevierStyleCrossRef" href="#fig0020">Fig&#46; 4</a>a&#44; <span class="elsevierStyleItalic">F</span>&#40;16&#44;<span class="elsevierStyleHsp" style=""></span>33&#41;<span class="elsevierStyleHsp" style=""></span>&#61;<span class="elsevierStyleHsp" style=""></span>7&#46;0769 <span class="elsevierStyleItalic">p</span><span class="elsevierStyleHsp" style=""></span>&#60;<span class="elsevierStyleHsp" style=""></span>0&#46;0001&#59; posterior probabilities are shown in Supplementary materials 2 and 3&#41;&#46; Linear discriminant factor from the DFA showed a contrasting distribution between the two groups of cases &#40;<a class="elsevierStyleCrossRef" href="#fig0020">Fig&#46; 4</a>&#41;&#46; The root factor was ordered mostly according to the following variables&#58; PrPD and AFL on the positive side and PrPA&#44; DFL and BLD on the negative side&#46;</p><elsevierMultimedia ident="fig0020"></elsevierMultimedia><p id="par0090" class="elsevierStylePara elsevierViewall">Results of the Mann&#8211;Whitney <span class="elsevierStyleItalic">U</span> test applied to meristic variables were also significant for three of five variables tested between species &#40;<a class="elsevierStyleCrossRef" href="#tbl0010">Table 2</a>&#41;&#44; and when applied to proportions&#44; comparisons were significant for 10 of 16 tested &#40;<a class="elsevierStyleCrossRef" href="#tbl0015">Table 3</a>&#41;&#46; Highly significant differences were found in the following proportions&#58; PrPA&#47;SL&#44; PrPP&#47;SL&#44; BD&#47;SL&#44; ADL&#47;SL&#44; BLD&#47;SL&#44; APL&#47;SL&#44; HL&#47;SL&#44; PrOL&#47;HL and ED&#47;SL&#46;</p><p id="par0095" class="elsevierStylePara elsevierViewall">Even when there is slight overlap&#44; significant differences were found between species in the following meristic variables &#40;<a class="elsevierStyleCrossRef" href="#tbl0015">Table 3</a>&#59; <span class="elsevierStyleItalic">p</span><span class="elsevierStyleHsp" style=""></span>&#60;<span class="elsevierStyleHsp" style=""></span>0&#46;001&#41;&#58; number of lateral scales&#44; with a mean of 33 and range of 30&#8211;35 in <span class="elsevierStyleItalic">P&#46; mixtlanensis</span>&#44; and 31 mean and range of 29&#8211;32 in <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#59; and with a lower number of anal-fin rays for <span class="elsevierStyleItalic">P&#46; oaxacae</span> &#40;12&#8211;14&#41;&#44; while for <span class="elsevierStyleItalic">P&#46; mixtlanensis</span> the anal-fin rays range from 12 to 21 &#40;<a class="elsevierStyleCrossRef" href="#fig0025">Fig&#46; 5</a>&#41;&#46; Given the previous results&#44; we conclude that <span class="elsevierStyleItalic">P&#46; mixtlanensis</span> fulfills the criteria to be considered a different taxonomic unit&#44; using both morphological and molecular &#40;<a class="elsevierStyleCrossRef" href="#bib0010">Doadrio et al&#46;&#44; 1999</a>&#41; criterion&#46;</p><elsevierMultimedia ident="fig0025"></elsevierMultimedia></span><span id="sec0045" class="elsevierStyleSection elsevierViewall"><span class="elsevierStyleSectionTitle" id="sect0065">Discussion</span><p id="par0100" class="elsevierStylePara elsevierViewall"><span class="elsevierStyleItalic">P&#46; mixtlanensis</span> and <span class="elsevierStyleItalic">P&#46; oaxacae</span> occur at geographically distant locations in the Mixteco basin&#44; and as we observed in the present description&#44; the two species are easily distinguishable by their body shapes and coloration patterns&#46; <span class="elsevierStyleItalic">P&#46; oaxacae</span> shows a characteristic spotty pattern along the lateral part of the body while in <span class="elsevierStyleItalic">P&#46; mixtlanensis</span> only shows a humeral spot&#46; The molecular distinctiveness of <span class="elsevierStyleItalic">P&#46; mixtlanensis</span> with respect to the other species within the <span class="elsevierStyleItalic">Profundulus</span> genus has been demonstrated by an allozyme study &#40;<a class="elsevierStyleCrossRef" href="#bib0010">Doadrio et al&#46;&#44; 1999</a>&#41;&#46; In addition&#44; phylogenetic analysis carried out by the authors showed that both&#44; <span class="elsevierStyleItalic">P&#46; oaxacae</span> and <span class="elsevierStyleItalic">P&#46; mixtlanensis</span> depict a high degree of divergence from the rest of the species within the subgenus <span class="elsevierStyleItalic">Profundulus</span> &#40;<span class="elsevierStyleItalic">Dp</span><span class="elsevierStyleHsp" style=""></span>&#61;<span class="elsevierStyleHsp" style=""></span>10&#46;5<span class="elsevierStyleHsp" style=""></span>&#177;<span class="elsevierStyleHsp" style=""></span>5&#46;9 mtDNA and <span class="elsevierStyleItalic">Dp</span><span class="elsevierStyleHsp" style=""></span>&#61;<span class="elsevierStyleHsp" style=""></span>1&#46;7<span class="elsevierStyleHsp" style=""></span>&#177;<span class="elsevierStyleHsp" style=""></span>0&#46;2 nDNA&#44; respectively&#44; data not shown&#41;&#46;</p></span></span>"
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          "titulo" => "Introduction"
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          "titulo" => "Materials and methods"
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            0 => array:2 [
              "identificador" => "sec0015"
              "titulo" => "Sample collection and taxonomic determination"
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              "identificador" => "sec0020"
              "titulo" => "Morphometric analyses"
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          "titulo" => "Description"
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              "identificador" => "sec0030"
              "titulo" => "Type locality&#58; Spring in Santiago Yosond&#250;a&#44; Oaxaca&#44; Mexico"
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            1 => array:2 [
              "identificador" => "sec0035"
              "titulo" => "Taxonomic summary"
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          "identificador" => "sec0040"
          "titulo" => "Results"
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        8 => array:2 [
          "identificador" => "sec0045"
          "titulo" => "Discussion"
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        9 => array:2 [
          "identificador" => "xack196110"
          "titulo" => "Acknowledgements"
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          "titulo" => "References"
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    "fechaRecibido" => "2014-08-13"
    "fechaAceptado" => "2015-07-22"
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            1 => "Killifish"
            2 => "Morphologic traits"
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            2 => "Caracteres morfol&#243;gicos"
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        "resumen" => "<span id="abst0005" class="elsevierStyleSection elsevierViewall"><p id="spar0005" class="elsevierStyleSimplePara elsevierViewall">The family Profundulidae is one of the few families of freshwater fishes endemic to the Mesoamerican region&#46; It includes the genus <span class="elsevierStyleItalic">Profundulus</span> that has been divided into two subgenera&#44; <span class="elsevierStyleItalic">Profundulus</span>&#44; with a wider distribution and <span class="elsevierStyleItalic">Tlaloc</span>&#44; which presents a more limited range&#46; The species currently recognized within the subgenus <span class="elsevierStyleItalic">Profundulus</span> are <span class="elsevierStyleItalic">P&#46; punctatus</span>&#44; <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#44; <span class="elsevierStyleItalic">P&#46; guatemalensis</span> and <span class="elsevierStyleItalic">P&#46; kreiseri</span>&#46; Results from a previous study using allozymes revealed a high degree of molecular differentiation of the populations inhabiting the Mixteca region &#40;located at the north of Oaxaca in Mexico&#41;&#44; compared to the rest of the species in the genus&#44; especially regarding its geographically nearest neighbor &#40;<span class="elsevierStyleItalic">P&#46; oaxacae</span>&#41;&#46; The main goal of this study was to evaluate the morphological distinctiveness and to assess the taxonomic identity of the populations from the Mixteca region in comparison with the morphological variation of <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#46; Based on 22 measurements &#40;17 morphometric and 5 meristic&#41;&#44; we examined the morphological variation within and among species&#44; including 50 individuals from 5 localities&#46; Our results show a high degree of morphological differentiation between both populations in accordance to the molecular information reported before&#44; supporting that populations from the Mixteca region are a new taxonomic entity&#44; which is described in the present study&#46;</p></span>"
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        "resumen" => "<span id="abst0010" class="elsevierStyleSection elsevierViewall"><p id="spar0010" class="elsevierStyleSimplePara elsevierViewall">La familia Profundulidae es una de las pocas familias de peces dulceacu&#237;colas end&#233;micas a la regi&#243;n de Mesoam&#233;rica&#46; La familia incluye al g&#233;nero <span class="elsevierStyleItalic">Profundulus</span>&#44; el cual a su vez incluye dos subg&#233;neros&#58; <span class="elsevierStyleItalic">Profundulus</span>&#44; con una mayor distribuci&#243;n y <span class="elsevierStyleItalic">Tlaloc</span>&#44; el cual presenta una distribuci&#243;n m&#225;s restringida&#46; Actualmente&#44; dentro del g&#233;nero <span class="elsevierStyleItalic">Profundulus</span> se incluyen las siguientes especies&#58; <span class="elsevierStyleItalic">P&#46; punctatus</span>&#44; <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#44; <span class="elsevierStyleItalic">P&#46; guatemalensis</span> y <span class="elsevierStyleItalic">P&#46; kreiseri</span>&#46; En un estudio previo basado en un an&#225;lisis de aloenzimas para el subg&#233;nero <span class="elsevierStyleItalic">Profundulus</span>&#44; se observ&#243; un alto grado de diferenciaci&#243;n entre las poblaciones de la regi&#243;n Mixteca&#44; al norte del estado de Oaxaca en M&#233;xico&#44; del resto de especies dentro del subg&#233;nero&#44; particularmente de las poblaciones de la especie geogr&#225;ficamente m&#225;s cercana &#40;<span class="elsevierStyleItalic">P&#46; oaxacae</span>&#41;&#46; El objetivo principal del presente estudio es caracterizar la variaci&#243;n morfol&#243;gica y evaluar la identidad taxon&#243;mica de las poblaciones de la regi&#243;n Mixteca compar&#225;ndolas con la variaci&#243;n morfol&#243;gica de <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#46; Esta caracterizaci&#243;n se llev&#243; a cabo con base en 22 variables &#40;17 morfom&#233;tricas y 5 mer&#237;sticas&#41; en 50 individuos provenientes de 5 localidades&#46; Nuestros resultados muestran que existe una clara diferenciaci&#243;n morfol&#243;gica entre ambos grupos de poblaciones&#44; en congruencia con evidencia molecular obtenida previamente&#44; y que sustenta el reconocimiento de las poblaciones de la regi&#243;n de la Mixteca como una nueva especie&#44; la cual es descrita en el presente estudio&#46;</p></span>"
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            "apendice" => "<p id="par0115" class="elsevierStylePara elsevierViewall">The following are the supplementary data to this article&#58;<elsevierMultimedia ident="upi0005"></elsevierMultimedia></p>"
            "etiqueta" => "Appendix A"
            "titulo" => "Supplementary data"
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          "en" => "<p id="spar0015" class="elsevierStyleSimplePara elsevierViewall">Map showing sampling points for the species&#58; <span class="elsevierStyleItalic">P&#46; mixtlanensis</span> sp&#46; nov&#46; &#40;black circles&#41; and <span class="elsevierStyleItalic">P&#46; oaxacae</span> &#40;white circles&#41;&#46;</p>"
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          "en" => "<p id="spar0020" class="elsevierStyleSimplePara elsevierViewall">Morphological comparison of the species of <span class="elsevierStyleItalic">Profundulus</span>&#58; &#40;a&#41; <span class="elsevierStyleItalic">Profundulus</span> &#40;<span class="elsevierStyleItalic">P&#46;</span>&#41; <span class="elsevierStyleItalic">mixtlanensis</span> sp&#46; nov&#46; and <span class="elsevierStyleItalic">P&#46; oaxacae</span> individuals&#59; &#40;b&#41; lateral view of the head of both species&#59; &#40;c&#41; dorsal head view of both species&#59; &#40;d&#41; holotype of <span class="elsevierStyleItalic">P&#46; mixtlanensis</span> sp&#46; nov&#46;</p>"
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          "en" => "<p id="spar0025" class="elsevierStyleSimplePara elsevierViewall">Bivariate plot of PCA results from morphometric measurements after allometric regression&#46; Solid circles correspond to <span class="elsevierStyleItalic">P&#46; mixtlanensis</span> sp&#46; nov&#46; and white to <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#46; PC1 explains 91&#46;73&#37; of the variance&#44; whereas PC2 explains 2&#46;79&#37; of the variance&#46;</p>"
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        "descripcion" => array:1 [
          "en" => "<p id="spar0030" class="elsevierStyleSimplePara elsevierViewall">Results from the discriminant function analysis &#40;DFA&#41;&#58; &#40;a&#41; proportions of correctly assigned individuals&#58; 97&#37; for <span class="elsevierStyleItalic">P&#46; mixtlanensis</span> sp&#46; nov&#46; &#40;individuals 1&#8211;35 in the graph&#41; and 100&#37; for <span class="elsevierStyleItalic">P&#46; oaxacae</span> &#40;individuals 36&#8211;50&#41;&#59; &#40;b&#41; distribution frequencies of root factor from the DFA between species&#46;</p>"
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          "en" => "<p id="spar0035" class="elsevierStyleSimplePara elsevierViewall">Frequency distribution of 5 meristic characters for <span class="elsevierStyleItalic">P&#46; mixtlanensis</span> sp&#46; nov&#46; &#40;black squares&#41; and <span class="elsevierStyleItalic">P&#46; oaxacae</span> &#40;white squares&#41;&#46;</p>"
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                  <table border="0" frame="\n
                  \t\t\t\t\tvoid\n
                  \t\t\t\t" class=""><thead title="thead"><tr title="table-row"><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">Map&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">Locality&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black"><span class="elsevierStyleItalic">N</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">Basin&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">Species proposed&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">Species by <a class="elsevierStyleCrossRefs" href="#bib0030">Miller &#40;1955&#44; 2005&#41;</a>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">UTM-X&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black">UTM-Y&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th></tr></thead><tbody title="tbody"><tr title="table-row"><td class="td" title="table-entry  " align="char" valign="top">1&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">R&#237;o Grande&#44; Santiago Juxtlahuaca&#44; Oaxaca&#44; M&#233;xico&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">5&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">Mixteco-Balsas&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; mixtlanensis</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; punctatus</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">605394&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">1910523&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry  " align="char" valign="top">2&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">Spring&#44; Sola de Vega&#44; Oaxaca&#46; M&#233;xico&#46;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">&#8211;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">Atoyac&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; oaxacae</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; oaxacae</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">715433&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">1826989&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry  " align="char" valign="top">3&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">R&#237;o Cucharas&#44; El Carmen Tuxtitl&#225;n&#44; Putla de Guerrero&#44; Oaxaca&#44; M&#233;xico&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">7&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">Atoyac&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; mixtlanensis</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; punctatus</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">615930&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">1884461&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry  " align="char" valign="top">4&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">R&#237;o Chiquito&#44; Tlaxiaco&#44; Oaxaca&#44; M&#233;xico&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">15&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">Mixteco-Balsas&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; oaxacae</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; oaxacae</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">643961&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">1905758&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry  " align="char" valign="top">5&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">Spring&#44; Aldama&#44; Oaxaca&#44; M&#233;xico&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">15&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">Atoyac&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; mixtlanensis</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; punctatus</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">648638&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">1875597&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry  " align="char" valign="top">6&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">Spring&#44; Chalcatongo&#44; Oaxaca&#44; M&#233;xico&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">&#8211;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">Atoyac&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; mixtlanensis</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; punctatus</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">650543&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">1884349&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry  " align="char" valign="top">7&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">Spring&#44; Santiago Yosond&#250;a&#44; Oaxaca&#46; M&#233;xico&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">8&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">Atoyac&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; mixtlanensis</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; punctatus</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">651751&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">1866154&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry  " align="char" valign="top">8&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">R&#237;o de las Grutas&#44; San Sebasti&#225;n de las Grutas&#44; Oaxaca&#44; M&#233;xico&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">&#8211;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">Atoyac&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; oaxacae</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top"><span class="elsevierStyleItalic">P&#46; oaxacae</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">718060&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">1839603&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr></tbody></table>
                  """
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          "en" => "<p id="spar0040" class="elsevierStyleSimplePara elsevierViewall">Sampling localities&#44; number of localities and number of individuals that were used in the morphologic analysis&#46; Taxonomic designation is according to our study&#44; and to <a class="elsevierStyleCrossRefs" href="#bib0030">Miller &#40;1955&#44; 2005&#41;</a>&#46;</p>"
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          "leyenda" => "<p id="spar0050" class="elsevierStyleSimplePara elsevierViewall">In bold highly significant <span class="elsevierStyleItalic">p</span>-values &#40;<span class="elsevierStyleItalic">p</span><span class="elsevierStyleHsp" style=""></span>&#60;<span class="elsevierStyleHsp" style=""></span>0&#46;005&#41; and in cursives significant <span class="elsevierStyleItalic">p</span>-values &#40;<span class="elsevierStyleItalic">p</span><span class="elsevierStyleHsp" style=""></span>&#60;<span class="elsevierStyleHsp" style=""></span>0&#46;05 and <span class="elsevierStyleItalic">p</span><span class="elsevierStyleHsp" style=""></span>&#62;<span class="elsevierStyleHsp" style=""></span>0&#46;005&#41;&#46;</p>"
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                0 => """
                  <table border="0" frame="\n
                  \t\t\t\t\tvoid\n
                  \t\t\t\t" class=""><thead title="thead"><tr title="table-row"><th class="td-with-role" title="table-head ; entry_with_role_rowhead " align="left" valign="top" scope="col">Morphometric measurements&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " colspan="2" align="center" valign="top" scope="col" style="border-bottom: 2px solid black">Mean &#40;standard error&#41;</th><th class="td" title="table-head  " colspan="4" align="center" valign="top" scope="col" style="border-bottom: 2px solid black">95&#37; Confidence interval</th><th class="td" title="table-head  " align="left" valign="top" scope="col"><span class="elsevierStyleItalic">F</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col"><span class="elsevierStyleItalic">p</span>-Value&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th></tr><tr title="table-row"><th class="td" title="table-head  " align="" valign="top" scope="col">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="" valign="top" scope="col">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="" valign="top" scope="col">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " colspan="2" align="center" valign="top" scope="col" style="border-bottom: 2px solid black">Upper limit</th><th class="td" title="table-head  " colspan="2" align="center" valign="top" scope="col" style="border-bottom: 2px solid black">Lower limit</th><th class="td" title="table-head  " align="" valign="top" scope="col">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="" valign="top" scope="col">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th></tr><tr title="table-row"><th class="td" title="table-head  " align="" valign="top" scope="col" style="border-bottom: 2px solid black">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black"><span class="elsevierStyleItalic">P&#46; mixtlanensis</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black"><span class="elsevierStyleItalic">P&#46; oaxacae</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black"><span class="elsevierStyleItalic">P&#46; mixtlanensis</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black"><span class="elsevierStyleItalic">P&#46; oaxacae</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black"><span class="elsevierStyleItalic">P&#46; mixtlanensis</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black"><span class="elsevierStyleItalic">P&#46; oaxacae</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="" valign="top" scope="col" style="border-bottom: 2px solid black">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="" valign="top" scope="col" style="border-bottom: 2px solid black">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th></tr></thead><tbody title="tbody"><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Standard length &#40;SL&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">63&#46;48 &#40;1&#46;70&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">47&#46;18 &#40;2&#46;41&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">66&#46;91&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">52&#46;04&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">60&#46;05&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">42&#46;33&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">30&#46;5&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Head length &#40;HL&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">17&#46;51 &#40;0&#46;52&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">13&#46;72 &#40;0&#46;74&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">18&#46;57&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">15&#46;22&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">16&#46;46&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">12&#46;22&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">17&#46;34&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pre-doral distance &#40;PrDD&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">42&#46;11 &#40;1&#46;11&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">31&#46;96 &#40;1&#46;58&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">44&#46;36&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">35&#46;15&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">39&#46;87&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">28&#46;78&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">27&#46;53&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pre-anal distance &#40;PrPA&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">42&#46;03 &#40;1&#46;09&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">32&#46;60 &#40;1&#46;54&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">44&#46;23&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">35&#46;72&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">39&#46;83&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">29&#46;49&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">24&#46;74&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Eye diameter &#40;ED&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">4&#46;15 &#40;0&#46;07&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">3&#46;41 &#40;0&#46;11&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">4&#46;31&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">3&#46;64&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">3&#46;99&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">3&#46;18&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">28&#46;63&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pre-orbital length &#40;PrOL&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">4&#46;80 &#40;0&#46;16&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">3&#46;36 &#40;0&#46;23&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">5&#46;13&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">3&#46;83&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">4&#46;47&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">2&#46;89&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">25&#46;35&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pre-pectoral distance &#40;PrPP&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">17&#46;77 &#40;0&#46;53&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">14&#46;08 &#40;0&#46;75&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">18&#46;84&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">15&#46;60&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">16&#46;70&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">12&#46;56&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">15&#46;98&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pre-ventral distance &#40;PrPV&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">32&#46;81 &#40;0&#46;93&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">25&#46;06 &#40;1&#46;32&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">34&#46;69&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">27&#46;73&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">30&#46;93&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">22&#46;39&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">22&#46;88&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Body depth &#40;BD&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">17&#46;46 &#40;0&#46;72&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">14&#46;70 &#40;1&#46;02&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">18&#46;91&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">16&#46;76&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">16&#46;00&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">12&#46;64&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">4&#46;85&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleItalic">0&#46;033</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Dorsal fin length &#40;DFL&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">10&#46;07 &#40;0&#46;32&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">7&#46;67 &#40;0&#46;46&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">10&#46;74&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">8&#46;60&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">9&#46;41&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">6&#46;73&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">17&#46;86&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pectoral-ventral length &#40;PVL&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">16&#46;74 &#40;0&#46;63&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">12&#46;54 &#40;0&#46;90&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">18&#46;02&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">14&#46;37&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">15&#46;45&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">10&#46;72&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">14&#46;28&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Anal fin length &#40;AFL&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">10&#46;33 &#40;0&#46;32&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">6&#46;95 &#40;0&#46;46&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">11&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">7&#46;89&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">9&#46;67&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">6&#46;02&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">35&#46;30&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pectoral fin length &#40;PFL&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">9&#46;76 &#40;0&#46;23&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">7&#46;19 &#40;0&#46;33&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">10&#46;23&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">7&#46;86&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">9&#46;29&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">6&#46;52&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">39&#46;77&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Ventral&#8211;anal length &#40;VAL&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">9&#46;59 &#40;0&#46;27&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">7&#46;75 &#40;0&#46;38&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">10&#46;13&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">8&#46;52&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">9&#46;04&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">6&#46;98&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">15&#46;34&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Body least depth &#40;BLD&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">8&#46;85 &#40;0&#46;28&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">7&#46;407 &#40;0&#46;40&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">9&#46;42&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">8&#46;21&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">8&#46;28&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">6&#46;59&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">8&#46;61&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;005</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Anal peduncle length &#40;APL&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">10&#46;53 &#40;0&#46;35&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">7&#46;024 &#40;0&#46;50&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">11&#46;25&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">8&#46;04&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">9&#46;81&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">6&#46;00&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">32&#46;22&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Caudal peduncle length &#40;CPL&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">10&#46;89 &#40;0&#46;33&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">8&#46;007 &#40;0&#46;46&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">11&#46;55&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">8&#46;94&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">10&#46;23&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">7&#46;06&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">25&#46;47&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td" title="table-entry  " colspan="9" align="left" valign="top"><span class="elsevierStyleVsp" style="height:0.5px"></span></td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Meristic counts&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="" valign="top">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="" valign="top">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " colspan="2" align="center" valign="top">Max</td><td class="td" title="table-entry  " colspan="2" align="center" valign="top">Min</td><td class="td" title="table-entry  " align="" valign="top">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="left" valign="top">Mann&#8211;Whitney <span class="elsevierStyleItalic">U</span> test&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Number of lateral scales &#40;E&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">33&#46;31 &#40;0&#46;20&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">30&#46;93&#40;0&#46;283&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">35&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">32&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">30&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">29&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="" valign="top">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Doral fin rays &#40;D&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">11&#46;45 &#40;0&#46;12&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">11&#46;4 &#40;0&#46;130&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">13&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">12&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">10&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">11&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="" valign="top">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;725&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Anal fin rays &#40;A&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">14&#46;45 &#40;0&#46;27&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">13&#46;13 &#40;0&#46;215&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">21&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">14&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">12&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">12&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="" valign="top">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;001</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Pectoral fin rays &#40;P&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">15&#46;94 &#40;0&#46;17&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">14&#46;33 &#40;0&#46;159&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">17&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">16&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">14&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">14&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="" valign="top">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">Ventral fin rays &#40;V&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">6&#46;05 &#40;0&#46;03&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">5&#46;93 &#40;0&#46;066&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">7&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">6&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">6&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">5&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="" valign="top">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;105&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr></tbody></table>
                  """
              ]
              "imagenFichero" => array:1 [
                0 => "xTab951138.png"
              ]
            ]
          ]
        ]
        "descripcion" => array:1 [
          "en" => "<p id="spar0045" class="elsevierStyleSimplePara elsevierViewall">Mean&#44; standard error and range values for morphometric measurements &#40;mm&#41; and meristic counts of <span class="elsevierStyleItalic">P&#46; mixtlanensis</span> and <span class="elsevierStyleItalic">P&#46; oaxacae</span>&#46; <span class="elsevierStyleItalic">p</span>-Values from the ANOVA test for morphometric measurements and those from the Mann&#8211;Whitney <span class="elsevierStyleItalic">U</span> test for meristic counts are provided&#46;</p>"
        ]
      ]
      7 => array:7 [
        "identificador" => "tbl0015"
        "etiqueta" => "Table 3"
        "tipo" => "MULTIMEDIATABLA"
        "mostrarFloat" => true
        "mostrarDisplay" => false
        "tabla" => array:2 [
          "leyenda" => "<p id="spar0060" class="elsevierStyleSimplePara elsevierViewall">SL&#44; standard length&#59; HL&#44; head length&#59; PrPA&#44; pre-anal distance&#59; PrPP&#44; pre-pectoral distance&#59; BD&#44; body depth&#59; AFL&#44; anal fin length&#59; VAL&#44; ventral&#8211;anal length&#59; BLD&#44; body least depth&#59; APL&#44; anal peduncle length&#59; PrOL&#44; pre-orbital length&#44; ED&#44; eye diameter&#46; In bold highly significant <span class="elsevierStyleItalic">p</span>-values &#40;<span class="elsevierStyleItalic">p</span><span class="elsevierStyleHsp" style=""></span>&#60;<span class="elsevierStyleHsp" style=""></span>0&#46;005&#41; and in cursives significant <span class="elsevierStyleItalic">p</span>-values &#40;<span class="elsevierStyleItalic">p</span><span class="elsevierStyleHsp" style=""></span>&#60;<span class="elsevierStyleHsp" style=""></span>0&#46;05 and <span class="elsevierStyleItalic">p</span><span class="elsevierStyleHsp" style=""></span>&#62;<span class="elsevierStyleHsp" style=""></span>0&#46;005&#41;&#46;</p>"
          "tablatextoimagen" => array:1 [
            0 => array:2 [
              "tabla" => array:1 [
                0 => """
                  <table border="0" frame="\n
                  \t\t\t\t\tvoid\n
                  \t\t\t\t" class=""><thead title="thead"><tr title="table-row"><th class="td-with-role" title="table-head ; entry_with_role_rowhead " align="left" valign="top" scope="col">Proportion&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col"><span class="elsevierStyleItalic">N</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " colspan="2" align="center" valign="top" scope="col" style="border-bottom: 2px solid black">Mean &#40;SE&#41;</th><th class="td" title="table-head  " colspan="2" align="center" valign="top" scope="col" style="border-bottom: 2px solid black">Upper limit</th><th class="td" title="table-head  " colspan="2" align="center" valign="top" scope="col" style="border-bottom: 2px solid black">Lower limit</th><th class="td" title="table-head  " align="left" valign="top" scope="col"><span class="elsevierStyleItalic">p</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th></tr><tr title="table-row"><th class="td" title="table-head  " align="" valign="top" scope="col" style="border-bottom: 2px solid black">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="" valign="top" scope="col" style="border-bottom: 2px solid black">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black"><span class="elsevierStyleItalic">P&#46; mixtlanensis</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black"><span class="elsevierStyleItalic">P&#46; oaxacae</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black"><span class="elsevierStyleItalic">P&#46; mixtlanensis</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black"><span class="elsevierStyleItalic">P&#46; oaxacae</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black"><span class="elsevierStyleItalic">P&#46; mixtlanensis</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="left" valign="top" scope="col" style="border-bottom: 2px solid black"><span class="elsevierStyleItalic">P&#46; oaxacae</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th><th class="td" title="table-head  " align="" valign="top" scope="col" style="border-bottom: 2px solid black">&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</th></tr></thead><tbody title="tbody"><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">PrPA&#47;SL&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">50&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;66 &#40;0&#46;0025&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;687 &#40;0&#46;0050&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;667&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;698&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;657&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;676&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">PrPP&#47;SL&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">50&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;28 &#40;0&#46;0040&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;302 &#40;0&#46;00376&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;289&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;310&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;272&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;294&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">BD&#47;SL&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">50&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;27 &#40;0&#46;006&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;310 &#40;0&#46;0043&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;285&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;320&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;260&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;301&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">AFL&#47;SL&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">50&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;16 &#40;0&#46;0025&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;151 &#40;0&#46;0039&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;169&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;159&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;159&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;142&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;011</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">VAL&#47;SL&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">50&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;15 &#40;0&#46;0021&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;163 &#40;0&#46;0048&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;155&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;173&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;146&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;153&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleItalic">0&#46;041</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">BLD&#47;SL&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">50&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;14 &#40;0&#46;0015&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;157 &#40;0&#46;0015&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;143&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;161&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;136&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;154&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;000</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">APL&#47;SL&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">50&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;16 &#40;0&#46;0024&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;148 &#40;0&#46;0032&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;166&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;155&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;156&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;142&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;001</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">HL&#47;SL&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">50&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;27 &#40;0&#46;0044&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;293 &#40;0&#46;0029&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;284&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;299&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;266&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;286&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;002</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">PrOL&#47;HL&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">50&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;28 &#40;0&#46;0103&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;244 &#40;0&#46;0071&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;305&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;259&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;263&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;228&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;007</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr><tr title="table-row"><td class="td-with-role" title="table-entry ; entry_with_role_rowhead " align="left" valign="top">ED&#47;SL&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">50&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;27 &#40;0&#46;0044&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;293 &#40;0&#46;0029&#41;&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;284&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;299&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;266&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top">0&#46;286&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td><td class="td" title="table-entry  " align="char" valign="top"><span class="elsevierStyleBold">0&#46;002</span>&nbsp;\t\t\t\t\t\t\n
                  \t\t\t\t</td></tr></tbody></table>
                  """
              ]
              "imagenFichero" => array:1 [
                0 => "xTab951136.png"
              ]
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          ]
        ]
        "descripcion" => array:1 [
          "en" => "<p id="spar0055" class="elsevierStyleSimplePara elsevierViewall">Mean&#44; standard error and range values for proportions of morphometric variables between <span class="elsevierStyleItalic">P&#46; oaxacae</span> and <span class="elsevierStyleItalic">P&#46; mixtlanensis</span>&#46; <span class="elsevierStyleItalic">p</span>-Values correspond to the Mann&#8211;Whitney <span class="elsevierStyleItalic">U</span> test&#46;</p>"
        ]
      ]
      8 => array:5 [
        "identificador" => "upi0005"
        "tipo" => "MULTIMEDIAECOMPONENTE"
        "mostrarFloat" => false
        "mostrarDisplay" => true
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Article information

ISSN: 18703453
Original language: English

DOI:10.1016/j.rmb.2015.10.001

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